For they can never truly die, they're killed day after day, going a little bit more insane, which creates. Many of these remaining humans were only civilians, but they would have time to learn, enough time, all the time in the world. Where all humans who died after its creation would revive, and be able to continue to search for a way to stop the dragonopolise. (Dragon endurance mode) Though essentially all the military was destroyed in this effort, they managed to save humanity, and made a lighthouse for lost souls. (This is endurance mode and hardcore mode, where the majority of the story takes place) To create this tower, many dragons were slain by RPGs among other weapons of destruction. In the midst of this combat and desolation, the military had a sudden breakthrough! Utilizing the bones of slain dragons, they created an immortality device of their own, one that would save both the occupant and their clothing, while unable to save anything not directly attached to the player's person. And thus have a hate of light, which reminds them of their days above ground. (Example, how many dragons are there at hell? Very few, showing their reverance of the dark lord.) The greator majority of skeltons can only arise underground, showing that they to have a connection to hell itself. The human skeltons are of a darker origin, they come from the same source as the dragons themselves, Satan. But the thirst for flesh will sometimes overcome this desire, and they will run straight through fire to catch their prey, and they will run off cliffs due to being blinded by bloodlust (See the portion of the game where they fall into the center of the earth via tracking you by a straight line).
The reason they hate light is that since they were all killed by the dragons, they remember the blinding flash before the end, reminding them of their pain, and humanity. Once killed by a dragon a person would become an immortal zombie, one that would regenerate after being killed, but has trouble regenerating through hard materials such as metal and brightly lit areas due to their hate of light. The military tried to halt their advance, but every person killed "Turned". Once in the age of man, cities spanned the Americas, then, the dragons came! They flew across the continent destroying cities and men, killing all who apposed them (This is evident by the amount of surface copper and stone, showing us that stone buildings fell and left behind scraps of metal across the continent due to wiring).
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You are now in a world full of flesh eating creatures who would like nothing more than to tear you apart.
However as you leave the blast doors shut behind you, there is no return. And you, the survivor, must leave the safehaven which has kept you safe for many years. However eventualy the oxygen inside starts running low.
The spawn tower is sealed off from the rest of the world and has its own oxygen supply and is the only man made structure to survive the initial "apocalypse". Reptilian creatures also were mutated by this, which effectivly became "Dragons". However these meteors contained hazardous gas which spread rapidly, turning any humans that we killed in the armagedon into mutant flesh eaters. This is "Infection" was caused after mysterious meteors started colliding with the earth which contained alien life forms. My own story / Theory ( If you can even call it that ) Is you are a survivor that made it to a nuclear bomb proof safehaven, The spawn tower, before a world engulfing infection spreads.